Harmonia vigintiduosignata

Harmonia vigintiduosignata

Harmonia vigintiduosignata is a Ladybird beetle (Coccinellidae) native to Africa.
Harmonia vigintiduosignata - TwentseWelle The other day I stumbled on a box with African beetles while doing some work in the museum TwentseWelle in Enschede (Netherlands) and it contained a Ladybird beetle I could not readily identify. Turns out it wasn't too hard: It looks a lot like a Harmonia and it has 22 spots, so when I found the name Harmonia vigintiduosignata for Africa somewhere that was a good lead :o)
I have only found two other images of this species on the internet, so I decided to augment that, even if these are just quick snapshots of an uncleaned museum specimen.
There is no date on the location tag, but the name Tanganyika suggests probably somewhere between 1961-65. Coccinellidae,Coccinellinae,Coleoptera,Harmonia,Harmonia vigintiduosignata,Ladybird,Tanzania,TwentseWelle

Appearance

It's a rather large Coccinellid (8.5–10.2 mm), shiny pale orange/yellowish brown with 22 spots, 11 per elytrum arranged in three cross bands of three spots each and an apical band of two (3,3,3,2).

References:

Some text fragments are auto parsed from Wikipedia.

Harmonia, vol.12 [cover photo by Julien Dabry]
http://archives.picardie-nature.org/?action=lecteur&document=876

Riaan Stals, Riaan; Goddy, Prinsloo (2007) Discovery of an alien invasive, predatory insect in South Africa: the multicoloured Asian ladybird beetle, Harmonia axyridis (Pallas) (Coleoptera: Coccinellidae) - South African Journal of Science, vol.103, pp.123-127.
[showing Harmonia vigintiduosignata on fig.4 pg.124]
http://www.scielo.org.za/pdf/sajs/v103n3-4/10.pdf

Old key here:
Mader, L. (1954) Coccinellidae - III. Teil. - Exploration du Parc National Albert - Exploratie van het Nationaal Albert Park - Mission/, vol.80, 195pp
Partial PDF for pages 94-195: http://biblio.naturalsciences.be/rbins-publications/institut-des-parcs-nationaux-du-congo/exploration-du-parc-national-albert-exploratie-van-het-nationaal-albert-park-mission-zending-g-f-de-witte-1933-1935/80-1954-mission-g-f-de-witte-coccinellidae/irscnb_p4137_rbins18799_80-98-210-part-2-red.pdf
Taxonomy
KingdomAnimalia
DivisionArthropoda
ClassInsecta
OrderColeoptera
FamilyCoccinellidae
GenusHarmonia
SpeciesHarmonia vigintiduosignata
Photographed in
Tanzania