Common Haircap Moss

Polytrichum commune

''Polytrichum commune'' is a species of moss found in many regions with high humidity and rainfall. The species can be exceptionally tall for a moss with stems often exceeding 30 cm though rarely reaching 70 cm, but it is most commonly found at shorter lengths of 5 to 10 cm.
Common Haircap Moss on Sphagnum Must have inspired the islands in the sky in "Avatar" Common Haircap Moss,Polytrichum commune,Wester Ross

Appearance

''Polytrichum commune'' is a medium to large moss. It is dark green in colour, but becomes brownish with age. The stems can occur in either loose or quite dense tufts, often forming extensive colonies. The stems are most typically found at lengths of 5 to 10 cm, but can be as short as 2 cm or as long as 70 cm.

They range in stiffness from erect to decumbent and are usually unbranched, though in rare cases they may be forked. The leaves occur densely to rather distantly, and bracts are present proximally.

The leaves typically measure 6 to 8 mm in length, but may be up to 12 mm long. When dry they are erect, but when moist they are sinuous with recurved tips and are generally spreading to broadly recurved, or sharply recurved from the base.

The leaf sheath is oblong to elliptic in outline, forming an involute tube and clasping the stem. This sheath is typically golden yellow and shiny, and it is abruptly contracted to the narrowly lanceolate blade.

Using a microscope, the marginal lamina can be seen to be level or erect, narrow, and typically 2 to 3 cells wide, though sometimes as many as 7 cells wide. It is toothed from the base of the blade up to the apex, with the teeth being unicellular and embedded in the margin. The costa, or central stalk of the leaf, is toothed on the underside near the apex, and is excurrent, meaning it extends beyond the end of the apex, ending in a short, rough awn.

The lamellae, ridges of cells that run along the leaf surface, are crenulate in profile and are 5 to 9 cells high. Their margins are distinctly grooved with 2 rows of paired, projecting knobs.

The marginal cells, when observed in section, may be narrow, but are more typically enlarged and wider than those beneath. They are retuse to deeply notched, and in rare cases are divided by a vertical partition. These cells are smooth and brownish in colour and have relatively thick cell walls. The sheath cells measure 60 to 90 µm long by 10 to 13 µm wide. These cells may be elongated rectangles or strongly linear structures up to 20 times long as wide. They become narrower toward the margins. Marginal lamina cells are 10 to 15 µm wide and are subquadrate .

The plants are sexually dioicous. The leaves of the perichaetium have a long sheath with a scarious margin, while the blades themselves are greatly reduced, gradually narrowing to a finely acuminate tip. These blades have toothed margins, are denticulate to subentire in outline, roughened to almost smooth, and have a costa that is excurrent.

The seta, or capsule stalk, is 5 to 9 cm long, and is stout and yellowish to reddish brown in colour. The capsule is 3 to 6 mm long, slightly rectangular to cubic in shape, and brown to dark reddish brown in colour. It is sharply 4 winged, inclined to horizontal, and glaucous when fresh. The peristome measures 250 µm, is pale in colour and has 64 teeth. The calyptra is golden yellow to brownish and completely envelops the capsule. The spores measure 5 to 8 µm, but may be up to 12 µm.

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Status: Unknown
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Taxonomy
KingdomPlantae
DivisionBryophyta
ClassPolytrichopsida
OrderPolytrichales
FamilyPolytrichaceae
GenusPolytrichum
SpeciesP. commune