AppearanceBody length: 9–13mm; wing length 10–12.5mm.
Head. Dark brown with gray pruinosity.
Antennae. Length 2.2–2.9 mm, reaching end of thorax to first abdominal segment. Scape and pedicel dark brown, first two flagellomeres yellow, next two yellow to light brown, remaining flagellomeres light brown.
Thorax. Mesonotum brown with largely gray pruinosity, margins without gray pruinosity and reddish brown. Brown longitudinal stripe across middle of pleura.
Wing. With generally four brown, transverse bands, interspersed with three clear, transverse bands; clear band at midlength and brown band following most distinct as bands. Transverse clear band across midlength of wing nearly always complete (band interrupted in some specimens, e.g., Hazleton Pennsylvania, Millville Newfoundland), bordering a fairly broad, recognizable, brown transverse band, brown band width from
just before end of Sc112 and extending through end of vein R1 at wing margin. Brown markings with obvious darker border clearly visible (‘‘ocellate’’), all markings fairly dark and well defined. Apex with distinct, irregular spots on apical margin from R3 to CuA1, sometimes connected as chain of markings. Distinct brown, circular or ocellus markings centered at origin of vein M (arculus area), origin of vein Rs, midlength of vein Cu (smaller and connected to Rs ocellus), surrounding discal cell and base of M1 and M2. Ocelli grown together in many specimens, clearly separated in others, total wing patterning variable.
Legs. Brownish yellow, femur with broad darker brown apical band (not subapical as in arizonense), some specimens with additional brown band near midlength blending gradually to yellow basally.
Abdomen. Tergites 1–7 yellowish brown, first two segments and posterior half of succeeding tergites with grayish pruinosity, lateral margins often darker brown, a median, longitudinal, brown band sometimes present; posterior margins of tergites narrowly pale. Sternites uniformly yellow. Apical margin with median fourth produced forward with shallow emargination between two slight lobes. (Apical ‘‘margin’’ in dry specimens most often straight because the lobes are often folded under and not observable.) Base of each interbase large, visible on lateral margins of tergite. Ninth sternite evenly sclerotized with conical protuberance medially.
Interbase. Very long and thin, extending past end of gonostyle, with a slight apical protuberance.
Proctiger. (observable in ethanol specimens) conical, with two small protuberances near apex, a pair of slightly sclerotized bands at midlength.
NamingEpiphragma fasciapenne (Say)
Limnobia fasciapennis Say, 1923
Limnophila pavonina Osten Sacken, 1859
FASCIAPENNIS: from the Latin "fascia" (a band) + "penna" (a wing, quill, feather); refers to the banded pattern on the wings of this species.
DistributionAlberta to Newfoundland, Canada, south to Kansas, Louisiana, and Florida, United States of America.
HabitatIn Kansas, the species was common in bottomland woods and around a swamp, with adults found on the ground or in vegetation less than 2 ft high. In Michigan, this species was also abundant in floodplain woods, swamps, and adjoining wooded ecotones. Larvae were found in sodden or submerged hardwood of all sizes and were able to survive partial desiccation of the wood.
Some text fragments are auto parsed from Wikipedia.https://bugguide.net/node/view/24964
Jon K. Gelhaus and Anthony Ruggeri. A review of the crane fly genus Epiphragma (Diptera: Tipulidae s.l.) in North America (including Mexico). Can. Entomol. 144: 353–375 (2012). DOI: 10.4039/tce.2012.32