Livelong Saxifrage

Appearance

"Saxifraga paniculata" is a perennial and stoloniferous herbaceous plant with flowering stems 10–30 cm in height. The most easily identifiable feature is its highly dense basal rosette of leaves, which are leathery, flat and stiff. 1–3 cm long, the oblong to ovate leaves are densely toothed and have fine leaf margins; a lime-encrusted white pore is present at the base of each leaf. The rosettes produce erect flowering stems, whilst the rosettes themselves grow at the end of runners.

The flowering stems have reduced and scattered leaves which terminate in a somewhat elongated cluster. The flowers are white, approximately 1 cm across and have dots which are either purplish or red. It flowers from mid-to-late June to early August, and produces perfect flowers. The flowers themselves have five petals, two styles, one inferior ovary and a two-beaked seed capsule. Warming noted that the flowers are protandrous, in the sense that even before the stigmas become receptive the flowers make and disperse pollen. However, as discussed in the 'Ecology' section below, the plant can self-pollinate.

"Saxifraga paniculata" can sometimes be mistaken for another plant in the same genus, "Saxifraga tricuspidata"; whilst "S. tricuspidata" does grow in a similar range, it can be differentiated from "S. paniculata" by its lack of lime-encrusted pores and by its crowded and much narrower leaves, which apart from three terminating stiff spine-tipped teeth have otherwise smooth margins.

Naming

"Saxifraga paniculata" was first formally described in the eighth edition of "The Gardeners Dictionary" by the Scottish botanist Philip Miller in 1768 and is placed in the genus "Saxifraga" and in the Saxifragaceae family. The generic name "Saxifraga" literally means "stone-breaker", from Latin ' + '. It is usually thought to indicate a medicinal use for treatment of urinary calculi, rather than breaking rocks apart.

Distribution

Growing in the Circumboreal Region, "Saxifraga paniculata" can be found throughout Central Europe, Greenland, Iceland and Scandinavia, as well as in the Caucasus and in North America where it can be found in the northern Great Lakes region, New England and New York State. Only historical records document it in Maine, whilst it is present, though considered a rarity, in Vermont, Nova Scotia, Newfoundland Island, New Brunswick, Minnesota and Labrador.

"Saxifraga paniculata" is a calciphile, and hence is found in calcareous habitats: an example of which is in a crevice of a basic rock. Of the rock crevices or rock ledges it does grow on, "S. paniculata" prefers the shady ones.

Habitat

Growing in the Circumboreal Region, "Saxifraga paniculata" can be found throughout Central Europe, Greenland, Iceland and Scandinavia, as well as in the Caucasus and in North America where it can be found in the northern Great Lakes region, New England and New York State. Only historical records document it in Maine, whilst it is present, though considered a rarity, in Vermont, Nova Scotia, Newfoundland Island, New Brunswick, Minnesota and Labrador.

"Saxifraga paniculata" is a calciphile, and hence is found in calcareous habitats: an example of which is in a crevice of a basic rock. Of the rock crevices or rock ledges it does grow on, "S. paniculata" prefers the shady ones.Its ability to close its leaf rosettes when undergoing deleterious environmental conditions such as excessive heat and droughts gives "Saxifraga paniculata" a very high resistance to sustained photoinhibition and irreversible dehydration; a paper published by Hacker and Neuner in 2006 found that "S. paniculata" was more resistant to cold induced photoinhibition in winter than any other evergreen subalpine species that the group had studied. Due to the short growing season and the possible lack of pollinators, "S. paniculata" can self-pollinate - whilst usually avoided in plant species as there is no potential for genetic variation in offspring, it does still ensure in dire conditions that seed is produced and dispersed.

In the wild, "Saxifraga paniculata" has been observed with many associated species, including but not limited to: "Trisetum spicatum", "Polygonum viviparum", "Polypodium virginianum", "Sagina nodosa", "Woodsia alpina", "Campanula rotundifolia", "Rubus pubescens", "Aralia nudicaulis," "Tortella tortuosa", "Aquilegia canadensis", "Carex eburnea" and "Woodsia glabella", as well as lichen cover.