Morchella snyderi

Appearance

It produces fruit bodies up to 14 cm tall with ridged and pitted conical caps, and stipes that become pitted in maturity. The color of the morel is yellow to tan when young, but the cap ridges become brown to black in maturity or when dried.

The fruit bodies are 6–14 cm high. The conical cap is 3.5–8 cm high and 3–5 cm wide at the widest point.

The cap surface features pits and ridges, formed by the intersection of 16–22 primary vertical ridges and frequent shorter, secondary vertical ridges, with occasional sunken, horizontal ridges. The cap is attached to the stipe with a sinus about 2–4 mm deep and 2–4 mm wide.

The ridges are smooth or very finely tomentose . They are initially pale yellowish, becoming pale tan, then grayish brown in maturity, eventually darkening to nearly black when dried. They are flattened when young but sometimes become sharpened or eroded in maturity.

The pits are somewhat elongated vertically . They are finely tomentose, yellowish when young, becoming pale tan to pale grayish brown. Fruit bodies are often found in a transitional stage where the upper part of the cap has turned dark while the lower part remains light.

The whitish to pale brownish stipe is 3.5–7 cm long by 2.5–4 cm wide and is roughly equal in width throughout its length, or sometimes slightly thicker near the base. Its surface, initially covered with whitish granules, becomes more granulated as the mushroom ages, and typically develops prominent ridges and pits with maturity.

The flesh is whitish, measuring 1–2 mm thick in the hollow cap; it becomes layered and chambered, particularly in the base of the stipe. The sterile inner surface of the cap is whitish and pubescent .

The ascospores of ''M. snyderi'' are elliptical and smooth, measuring 25–37 by 15–23 µm. Asci are eight-spored, cylindrical, and measure 225–300 by 7.5–32.5 µm. Paraphyses are cylindrical, septate, and measure 100–200 by 7.5–20 µm. Their tips are variably shaped, from rounded to club-shaped, to fuse-shaped.

The contents of the paraphyses are hyaline to faintly brownish in dilute potassium hydroxide. Hyphae on the sterile cap ridges are septate and measure 75–175 by 10–20 µm. The terminal cells are variably shaped, and have hyaline to brownish contents in KOH.

Naming

''Morchella snyderi'' can be distinguished from similar North American morels by differences in ecological and morphological features. ''M. frustrata'' resembles young specimens of ''M. snyderi'', but the ridges of the caps of the former species do not darken in maturity, and it has smaller ascospores, measuring 20–29 by 14–19 µm. Another potential lookalike, ''M. brunnea'', has a browner fruit body color in young specimens, and a stipe that is not lacunose.

Distribution

Fruiting occurs from April to June. The fungus has been collected in California, Idaho, Montana, Oregon, Washington, New Mexico, and Arizona. ''M. snyderi'', identified as phylogenetic species ''Mel''-12, has been shown to colonize the non-native species ''Bromus tectorum'' as an endophyte. This is hypothesized to be a contributing factor in the success of cheatgrass as an invasive species in western North America.

Habitat

''Morchella snyderi'' is suspected of being both saprobic and mycorrhizal at different stages in its life cycle. Fruit bodies grow singly, scattered, or in groups on the ground under conifers, particularly Douglas fir, ponderosa pine and white fir.