Juniper haircap moss

Polytrichum juniperinum

''Polytrichum juniperinum'', commonly known as juniper haircap moss, is an evergreen and perennial species of moss that is widely distributed, growing on every continent including Antarctica.
A Forest of Sporophytes Waiting for the sun to dry these Polytrichum juniperinum sporophytes so the spores can be released.
 Canada,Geotagged,Juniper haircap moss,Polytrichum juniperinum,Spring

Appearance

The stems are reddish with grey-green leaves that have a distinctive red-brown tip. This characteristic allows them to be separated from the bristly haircap, a plant that the juniper haircap moss have a close resemblance to; the difference is that the bristly haircap have a green tip. The leaves of juniper haircap moss are lanceolate and upright spreading when dry, and when moist, wide-spreading. Although their growth form can be varied, they generally grow in thin, interwoven mats, and hardly as closely associated individuals. Juniper haircap moss have a well-developed system of tiny tubes for carrying water from the rhizoids to leaves that is uncharacteristic of mosses, resembling the system that has evolved in vascular plants such as ferns, gymnosperms and angiosperms. As a result of this developed system, stems have greater potential for height than in typical mosses.
A Sporangium Revealed. This series of photos shows the sporangium revealed. After removing the fibrous calyptra you can see the elf cap shaped operculum. Upon removal of the operculum the really interesting structure, to me, is revealed. The periostome teeth surround the opening of the sporangium and the opening is sealed by a membranous epiphragm. When the humidity decreases the epiphragm ruptures and the peristome teeth regulate the dispersal of the spores from the sporangium. When the humidity increases the teeth close the peristome keeping the spores safe inside the sporangium, when the humidity decreases the teeth bend outward allowing the spores to be dispersed by the dry breezes. What allows the teeth to act this way is that the inside and the outside surfaces of the teeth are structurally different and therefore respond differently to humidity.
There is a good micro photograph of the cross section of a similar sporangium on this webpage.
http://www3.botany.ubc.ca/bryophyte/poly1.htm Canada,Geotagged,Juniper haircap moss,Polytrichum juniperinum

Distribution

Juniper haircap moss grow across a wide gradient of habitats but it is most commonly found on dry, acidic, exposed habitats. It is frequent in areas that previously experienced disturbances such as fire and logging. Other areas they occupy are mineral soil, humus and rocks, stumps, banks, trailsides and dry open woods. Although Juniper haircap moss is not usually found in moist or wet environments, it has been found growing on moist woods and other moist sites such as streambanks.
Dewy Juniper Haircap Moss Polytrichum juniperinum is distinguished from P. commune and P. piliferum by its red hair points on the leaves. P. commune has none but toothed margins on the leaves while P. piliferum has white hairpoints on the leaves. The flower like structures are the male “flowers” (antherdial heads)  which produce the sperm.
For the life cycle have a look here, https://commons.m.wikimedia.org/wiki/File:Moss_life_Cycle.jpg Canada,Geotagged,Juniper haircap moss,Polytrichum juniperinum,Spring

Reproduction

It is a dioecious plant, meaning that the male and female gametophytes are on separate plants. Juniper haircap moss have very obvious male and female parts. Male plants are said to be unusual because they continue growing without losing the old male organs. The male plants are very noticeable due to their bright reddish orange modified leaves that form small terminal 'flowers' at the shoot ends. The plant has a gametophyte dominant life cycle similar to other mosses. Water is required for reproduction to take place, to enable the sperm to swim down the neck of the archegonia to reach the egg. Once fertilization takes place, the sporophyte of the juniper haircap moss lives on the female gametophyte, growing out of the archegonia. The sporophyte consists of a foot, stalk, a spore capsule, an operculum, and a calyptra. There are 64 short blunt teeth at the top surrounding the capsule mouth and the hood of the capsule, the calyptra, has long hairs that extends down the entire length of the capsule, hence the name 'haircap moss'.

References:

Some text fragments are auto parsed from Wikipedia.

Status: Unknown
EX EW CR EN VU NT LC
Taxonomy
KingdomPlantae
DivisionBryophyta
ClassPolytrichopsida
OrderPolytrichales
FamilyPolytrichaceae
GenusPolytrichum
Species