Cypripedium calceolus

Appearance

This is the largest orchid species in Europe, growing to 60 cm tall with flowers as wide as 9 cm. Before it flowers, it is distinguished from other orchids by the large size and width of its ovate leaves , which like other orchids exhibit parallel venation. Each shoot has up to four leaves and a small number of flowers, which have long often twisted petals varying from red-brown to black and a slipper-shaped yellow labellum, within which red dots are visible. It is a long-lived perennial and spreads using horizontal stems .

''Cypripedium calceolus'' can be confused, when not flowering, with ''Allium ursinum'', ''Convallaria majalis'' or several species of ''Epipactis'' orchid. It closely resembles other species of ''Cypripedium'' orchid native to the United States .

Chromosome number has been given as 2n=20 but also 2n=22

Distribution

It has a widespread distribution from Europe east through Asia from Spain to the Pacific, including almost every country in Europe plus Russia , northeastern China , Mongolia, Korea and Rebun Island in Japan.

It is typically found in open woodland on moist calcareous soils. In continental Europe it is also found growing in the decomposed humus of semi-shaded woodland cover on limestone. It has declined over much of the European part of its range, and as a result is legally protected in a number of countries. Its upper elevation limit is 2100m. According to growing instructions provided by Phytesia, it cannot survive waterlogging or direct midday sun.

Status

Although the global conservation status of ''Cypripedium calceolus'' is least concern according to the IUCN Red List, in many countries it has become rare and is afforded legal protection. ''C. calceolus'' is common in Poland and Austria but in Greece it has become extinct.

In Britain, it was formerly a reasonably widespread plant across northern England, particularly the limestone area of the Yorkshire Dales. By the late 20th century it had declined to just a single plant in one location in the dales. While the virtual extinction of the lady's-slipper orchid from its historical range in Britain is often blamed on uprooting by gardeners and botanists, it is also the case that its preferred habitat shrank markedly with human clearance of woodland from the limestone landscape, and the grazing of sheep will have finished it off. It became a protected species in the UK in 1975 under the Conservation of Wild Creatures and Wild Plants Act, but a reintroduction programme for the lady's-slipper orchid is in place, and has led to a population of hundreds of plants as of 2003.

A laboratory in Belgium has developed a technique to mass-produce ''C. calceolus'', so plants are commercially available, unlike many orchid species.

Habitat

It has a widespread distribution from Europe east through Asia from Spain to the Pacific, including almost every country in Europe plus Russia , northeastern China , Mongolia, Korea and Rebun Island in Japan.

It is typically found in open woodland on moist calcareous soils. In continental Europe it is also found growing in the decomposed humus of semi-shaded woodland cover on limestone. It has declined over much of the European part of its range, and as a result is legally protected in a number of countries. Its upper elevation limit is 2100m. According to growing instructions provided by Phytesia, it cannot survive waterlogging or direct midday sun.''Cypripedium calceolus'' is frequently associated with stands of hazel trees.

It is pollinated by a number of different insect species, including at least seven species of miner bee in the genera ''Andrena'' and ''Colletes'', as well as at least two species of ''Lasioglossum'' . Plants are very vulnerable to herbivory by slugs and snails.

''Cypripedium calceolus'' is known to primarily associate with mycorrhizal fungi in the genus ''Tulasnellaceae''. Specific relationships with mycorrhizal fungi are key to orchids' ability to access soil nutrients. Other suggested mycorrhizal partners include ''Alternaria'' sp., ''Ceratorhiza'' sp., ''Chaetomium'' sp., ''Cylindrocarpon'' sp., ''Epicoccum purpureum'', ''Epulorhiza'' sp., ''Moniliopsis'' sp., ''Mycelium radicis atrovirens'', ''Phoma'' sp. and ''Rhizoctonia subtilis'' sp.

Cultural

''Cypripedium calceolus'' has appeared on postage stamps in a huge number of countries including Austria, the Czech Republic, Denmark, France, Grenadines, Hungary, Italy, Latvia, Madagascar, Moldova, Mozambique, Norway, Romania, Russia, Slovenia, Sweden, Uganda, Ukraine and the United Kingdom.

The Norwegian municipality of Snåsa has a ''Cypripedium calceolus'' in its coat-of-arms.

In Pavel Ivanovich Melnikov's "In the Forests", a ''znakharka'' calls this Adam's head, Adam's grass, and Cuckoo's slippers and says it is good for every ill including driving away evil spirits.