Brown-eyed Camouflage Lichen

Thallus: adnate to loosely adnate appressed throughout or somewhat raised at the periphery, foliose, up to 8 (-11) cm diam., lobate lobes: short and rounded, ± contiguous to imbricate, (1-) 2-6 (-l 0) mm broad, ± flat upper surface: olive-brown to dark brown, smooth to weakly rugose or pitted near the periphery, inward becoming more strongly rugose and often grossly so, sometimes with isidioid papillae or tubercles, these verging on true isidia in extreme cases; dull throughout or somewhat shiny on the lobe-ends; pseudocyphellae usually sparse or absent, occasionally rather numerous but even then small and rather obscure, the laminal tubercles developed by some specimens often bearing distinctive pseudocyphellae lower surface: dark brown or black, pale brown to tan at the periphery or occasionally throughout; smooth to rugulose, dull to rather shiny; moderately to rather densely rhizinate, the rhizines concolorous with the lower surface or paler Apothecia: common, up to 5 mm diam., sessile to short stipitate, concave to flattening or sometimes convex, the margin entire to more often crenate or tuberculate, often with rather numerous pseudocyphellae, especially on the tubercles asci: clavate, 8-spored ascospores: ellipsoid to subglobose, (7-) 8-10.5 x 5-8 (-9) µm Pycnidia: common, immersed conidia: acerose to weakly fusiform or weakly bifusiform, 6.5-8.5 (-10) x 1 µm Spot tests: cortex K-, C-, KC-, P-, HNO3-; medulla K-, C-, KC-, P- Secondary metabolites: none detected.